:root{--forest:#173126;--paper:#f4f0e7;--paper-deep:#e9e2d5;--gold:#9a8142;--muted:#687168;--line:rgba(23,49,38,.15)}
*{box-sizing:border-box;margin:0;padding:0}html{scroll-behavior:smooth}body{background:var(--paper);color:var(--forest);font-family:Arial,Helvetica,sans-serif}a{color:inherit}.bio-header{position:relative;z-index:20;display:flex;align-items:center;justify-content:space-between;min-height:94px;padding:8px 5.5%;border-bottom:1px solid var(--line);background:rgba(244,240,231,.94);backdrop-filter:blur(12px)}.bio-brand{display:block;width:clamp(310px,38vw,470px);line-height:0}.bio-brand img{display:block;width:100%;max-height:78px;object-fit:contain;object-position:left center}.bio-nav{display:flex;align-items:center;gap:29px;font-size:12px}.bio-nav a{text-decoration:none}.bio-nav a:not(.back-link){color:#46544a}.bio-nav a:not(.back-link):hover{color:var(--gold)}.back-link{display:inline-flex;align-items:center;gap:10px;padding:12px 17px;border:1px solid rgba(23,49,38,.23);border-radius:30px;font-size:10px;font-weight:700;letter-spacing:.12em;text-transform:uppercase;transition:.3s}.back-link:hover{background:var(--forest);color:var(--paper)}
.bio-hero{position:relative;overflow:hidden;padding:110px 7% 105px;background:var(--paper-deep);isolation:isolate}.bio-hero:before{content:"";position:absolute;z-index:-1;right:-210px;top:-300px;width:650px;height:650px;border:1px solid rgba(154,129,66,.17);border-radius:50%;box-shadow:0 0 0 78px rgba(154,129,66,.03),0 0 0 156px rgba(154,129,66,.02)}.bio-hero-inner{max-width:1240px;margin:auto}.eyebrow{display:flex;align-items:center;gap:13px;margin-bottom:23px;color:var(--gold);font-size:11px;font-weight:700;letter-spacing:.21em;text-transform:uppercase}.eyebrow:before{content:"";width:34px;height:1px;background:var(--gold)}.bio-hero h1{max-width:920px;font:400 clamp(3.5rem,7vw,7rem)/.96 Georgia,serif;letter-spacing:-.045em}.bio-hero-copy{display:grid;grid-template-columns:minmax(0,1fr) minmax(320px,.52fr);gap:70px;align-items:end;margin-top:44px}.bio-hero-copy p{max-width:640px;color:#536158;font:400 clamp(1.05rem,1.6vw,1.3rem)/1.75 Georgia,serif}.bio-hero-meta{padding-left:26px;border-left:1px solid rgba(154,129,66,.45);color:var(--muted);font-size:11px;line-height:1.8;letter-spacing:.08em;text-transform:uppercase}
.category-links{display:grid;grid-template-columns:repeat(3,1fr);max-width:1240px;margin:-42px auto 0;padding:0 7%;position:relative;z-index:3;gap:15px}.category-link{position:relative;min-height:360px;padding:34px;border-radius:21px;background:var(--forest);color:var(--paper);overflow:hidden;text-decoration:none;box-shadow:0 24px 55px rgba(23,49,38,.14);transition:transform .45s ease,box-shadow .45s ease}.category-link:hover{transform:translateY(-8px);box-shadow:0 34px 70px rgba(23,49,38,.2)}.category-link:before{content:"";position:absolute;right:-80px;top:-85px;width:220px;height:220px;border:1px solid rgba(193,168,102,.25);border-radius:50%;box-shadow:0 0 0 38px rgba(193,168,102,.035)}.category-no{position:relative;color:#c1a866;font:400 3.6rem/1 Georgia,serif}.category-link-copy{position:absolute;left:34px;right:34px;bottom:31px}.category-link h2{font:400 clamp(1.8rem,3vw,2.65rem)/1.05 Georgia,serif}.category-link p{margin-top:13px;color:rgba(244,240,231,.65);font-size:12px;line-height:1.65}.category-link-footer{display:flex;align-items:center;justify-content:space-between;margin-top:24px;padding-top:18px;border-top:1px solid rgba(244,240,231,.16);color:#d3bd79;font-size:9px;font-weight:700;letter-spacing:.14em;text-transform:uppercase}.category-arrow{font-size:20px;transition:transform .3s}.category-link:hover .category-arrow{transform:translateX(6px)}
.bio-intro{max-width:1240px;margin:auto;padding:104px 7% 68px}.bio-intro-grid{display:grid;grid-template-columns:minmax(0,.85fr) minmax(0,1.15fr);gap:80px;align-items:start}.bio-intro h2{font:400 clamp(2.7rem,5vw,4.6rem)/1.02 Georgia,serif;letter-spacing:-.03em}.bio-intro p{color:var(--muted);font-size:14px;line-height:1.9}.taxonomy-note{margin-top:22px;padding-top:20px;border-top:1px solid var(--line);color:#7c735e!important;font-size:11px!important}
.catalogue{max-width:1240px;margin:auto;padding:0 7% 110px}.catalogue-tools{display:flex;align-items:center;justify-content:space-between;gap:30px;margin-bottom:31px;padding-bottom:22px;border-bottom:1px solid var(--line)}.species-count{font:400 1.3rem Georgia,serif}.species-count strong{color:var(--gold);font-size:2rem;font-weight:400}.search-wrap{position:relative;width:min(100%,340px)}.species-search{width:100%;padding:13px 44px 13px 17px;border:1px solid rgba(23,49,38,.2);border-radius:30px;background:rgba(255,255,255,.32);color:var(--forest);outline:none}.species-search:focus{border-color:var(--gold);background:#fff}.search-wrap:after{content:"⌕";position:absolute;right:17px;top:8px;color:var(--gold);font-size:22px}
.species-grid{display:grid;grid-template-columns:repeat(3,minmax(0,1fr));gap:22px}.species-card{position:relative;min-height:455px;border-radius:20px;overflow:hidden;background:var(--forest);color:#fff;box-shadow:0 20px 48px rgba(23,49,38,.12);opacity:0;transform:translateY(25px);transition:opacity .75s ease,transform .75s cubic-bezier(.2,.75,.25,1),box-shadow .4s ease}.species-card.visible{opacity:1;transform:none}.species-card:hover{box-shadow:0 30px 62px rgba(23,49,38,.22)}.species-image{position:absolute;inset:0;background:linear-gradient(145deg,#294a3a,#173126);overflow:hidden}.species-image:before{content:attr(data-initial);position:absolute;left:50%;top:42%;transform:translate(-50%,-50%);color:rgba(244,240,231,.1);font:400 8rem Georgia,serif}.species-image img{width:100%;height:100%;object-fit:cover;opacity:0;transform:scale(1.04);filter:saturate(.86);transition:opacity .6s ease,transform .9s ease}.species-image img.loaded{opacity:1}.species-card:hover img{transform:scale(1.09)}.species-card:after{content:"";position:absolute;inset:0;background:linear-gradient(180deg,rgba(7,21,14,.03) 27%,rgba(8,24,16,.9) 82%,rgba(8,24,16,.98))}.status-tag{position:absolute;z-index:2;top:20px;right:20px;padding:9px 11px;border:1px solid rgba(255,255,255,.34);border-radius:30px;background:rgba(23,49,38,.76);backdrop-filter:blur(7px);color:#f1dc9d;font-size:8px;font-weight:700;letter-spacing:.12em;text-transform:uppercase}.species-info{position:absolute;z-index:2;left:27px;right:27px;bottom:25px}.species-index{display:block;margin-bottom:10px;color:#d3bd79;font-size:9px;font-weight:700;letter-spacing:.16em}.species-info h2{font:400 2rem/1.05 Georgia,serif}.species-info em{display:block;margin-top:8px;color:rgba(255,255,255,.7);font:italic 13px Georgia,serif}.species-group{display:block;margin-top:17px;padding-top:15px;border-top:1px solid rgba(255,255,255,.2);color:rgba(255,255,255,.62);font-size:9px;font-weight:700;letter-spacing:.12em;text-transform:uppercase}.no-results{display:none;padding:60px 0;color:var(--muted);font:400 1.5rem Georgia,serif}.no-results.show{display:block}
.historic-note{max-width:1240px;margin:0 auto 105px;padding:0 7%}.historic-note-inner{display:grid;grid-template-columns:120px 1fr;gap:34px;align-items:center;padding:34px 38px;border-top:1px solid rgba(154,129,66,.48);border-bottom:1px solid rgba(154,129,66,.48)}.historic-number{color:var(--gold);font:400 3rem/1 Georgia,serif}.historic-note h2{font:400 1.7rem/1.15 Georgia,serif;margin-bottom:8px}.historic-note p{color:var(--muted);font-size:12px;line-height:1.7}
.bio-footer{padding:48px 7%;background:var(--forest);color:var(--paper)}.bio-footer-inner{max-width:1240px;margin:auto;display:flex;align-items:center;justify-content:space-between;gap:30px}.bio-footer h2{font:400 1.45rem Georgia,serif}.bio-footer p{margin-top:8px;color:rgba(244,240,231,.58);font-size:10px;letter-spacing:.08em}.bio-footer a{color:#d3bd79;font-size:10px;letter-spacing:.12em;text-decoration:none;text-transform:uppercase}
@media(max-width:950px){.bio-nav a:not(.back-link){display:none}.bio-hero-copy,.bio-intro-grid{grid-template-columns:1fr;gap:28px}.category-links,.species-grid{grid-template-columns:repeat(2,1fr)}.category-link:last-child{grid-column:1/-1}.species-grid .species-card:last-child:nth-child(odd){grid-column:1/-1}.species-grid .species-card:last-child:nth-child(odd){max-width:calc(50% - 11px)}}
@media(max-width:650px){.bio-header{min-height:78px;padding:8px 18px}.bio-brand{width:230px}.back-link{padding:10px 12px;font-size:8px}.bio-hero{padding:80px 20px 82px}.bio-hero h1{font-size:3.35rem}.category-links{grid-template-columns:1fr;margin-top:-30px;padding:0 20px}.category-link,.category-link:last-child{grid-column:auto;min-height:310px}.bio-intro{padding:80px 20px 50px}.catalogue{padding:0 20px 80px}.catalogue-tools{align-items:flex-start;flex-direction:column}.search-wrap{width:100%}.species-grid{grid-template-columns:1fr}.species-grid .species-card:last-child:nth-child(odd){grid-column:auto;max-width:none}.species-card{min-height:430px}.historic-note{padding:0 20px;margin-bottom:75px}.historic-note-inner{grid-template-columns:1fr;padding:29px 4px}.bio-footer-inner{display:block}.bio-footer a{display:inline-block;margin-top:22px}}
@media(prefers-reduced-motion:reduce){html{scroll-behavior:auto}.category-link,.species-card,.species-image img{transition:none}.species-card{opacity:1;transform:none}}

/* Palette lock: reuse only the established website colours. */
.bio-nav a:not(.back-link),.bio-hero-copy p,.taxonomy-note{color:var(--muted)!important}
.species-image{background:var(--forest)}
.species-card:after{background:linear-gradient(180deg,rgba(23,49,38,.03) 27%,rgba(23,49,38,.9) 82%,rgba(23,49,38,.98))}
.status-tag{color:#d3bd79}